# vim: ts=50 nowrap noet textwidth=0
#
#  Title: Phylogeny of Ferns and Fern Allies
#  Label: Ferns
#  Strict: 1
#
#  Notes: <p>This is based largely on Pryer et al 1995, with a few
#  modifications from other stuff I found on the web.  I got the number of
#  genera and species, and the placement of genera came from <a
#  href="http://homepages.caverock.net.nz/~bj/fern">Checklist of World
#  Ferns</a>.
#
#  Link: top            top.html?node=Filicophyta
#  Link: Spermatophyta  top.html?node=Spermatophyta
#
################################################################################

* Embryophyta
|* Lycophyta
||. Isoet{ales,aceae,es}
||' Selaginell{ales,aceae,a}
|' Lyc{opsida,opodales,opodaceae}
|.* Filicophyta
||| Equiset{opsida,ales,aceae,um}
||' Filicopsida
'' Spermatophyta

* Filicopsida
| Psilot{ales,aceae}
|. Maratti{ales,aceae}
|' Ophiogloss{ales,aceae}
|* Leptosporangiate_Ferns
|| Hymenophyllopsidaceae
|| Osmundaceae
||. Gleicheniaceae
||| Hymenophyllaceae
|||. Matoniaceae
||||. Cheiropleuriaceae
||||' Dipteridaceae
||||. Schizae{ales,aceae}
|||||* Marsileales
|||||| Marsileaceae
||||||. Azollaceae
|||||'' Salviniaceae
|||||* Cyatheales
|||||| Plagiogyriaceae
||||||. Cyatheaceae
||||||| Dicksoniaceae
||||||| Lophosoriaceae
||||||| Loxsomataceae
|||||'' Metaxyaceae
|||||* Polypodiaceous_Ferns
|||||| Lindsae{ales,aceae}
||||||. Dennstaedti{ales,aceae}
|||||||* Pteridoid_Ferns
||||||||. Pteridaceae
|||||||'' Vittariaceae
|||||||. Aspleni{ales,aceae}
|||||||| Blechnaceae
|||||||| Thelypteridaceae
||||||||. Dryopteridaceae
|||||||||. Nephrolepidaceae
||||||||||. Davalliaceae
|||||||||||. Grammitidaceae
'''''''''''' Polypodiaceae

* Aspleniaceae
| Asplenium
| Camptosorus
| Diellia
| Holodictyum
| Pleurosorus
' Schaffneria

* Azollaceae
' Azolla

* Blechnaceae
| Blechnum
| Brainea
| Doodia
| Pteridoblechnum
| Sadleria
| Salpichlaena
| Stenochlaena
' Woodwardia

* Cheiropleuriaceae
' Cheiropleuria

* Cyatheaceae
| Alsophila
| Cnemidaria
| Cyathea
' Sphaeropteris

* Davalliaceae
| Araiostegia
| Davallia
| Davallodes
| Gymnogrammitis
' Leucostegia

* Dennstaedtiaceae	(subfamilies are old families, may not be monophyletic)
|* Dennstaedtioideae
|| Coptidipteris
|| Costaricia
|| Dennstaedtia
|| Leptolepia
|| Microlepia
|' Oenotrichia
|* Hypolepidoideae
|| Blotiella
|| Histiopteris
|| Hypolepis
|| Paesia
|' Pteridium
|* Monachosoroideae
'' Monachosorum

* Dicksoniaceae
| Calochlaena
| Cibotium
| Culcita
| Cystodiopteris
| Dicksonia
' Thyrsopteris

* Dipteridaceae
' Dipteris

* Gleicheniaceae
| Dicranopteris
| Diplopterygium
| Gleichenia
| Sticherus
' Stromatopteris

* Grammitidaceae
| Acrosorus
| Adenophorus
| Calymmodon
| Ceradenia
| Chrysogrammitis
| Cochlidium
| Ctenopteris
| Enterosora
| Grammitis
| Lellingeria
| Melpomene
| Micropolypodium
| Prosaptia
| Scleroglossum
| Terpsichore
| Themelium
| Xiphopteris
' Zygophlebia

* Hymenophyllaceae	(subfamilies are old families, may not be monophyletic)
|* Hymenophylloideae
|| Cardiomanes
|| Craspedophyllum
|| Hymenoglossum
|| Hymenophyllum
|' Serpyllopsis
|* Trichomanoideae
|| Callistopteris
|| Cephalomanes
|| Crepidomanes
|| Gonocormus
|| Macroglena
|| Microtrichomanes
|| Nesopteris
|| Pleuromanes
|| Reediella
|| Selenodesmium
'' Trichomanes

* Hymenophyllopsidaceae
' Hymenophyllopsis

* Lindsaeaceae	(subfamilies are old families, may not be monophyletic)
|* Lindsaeoideae
|| Lindsaea
|| Ormoloma
|| Tapeinidium
|' Xyropteris
|* Lonchitoideae
|| Lonchitus
|| Odontosoria
'' Saccoloma

* Lophosoriaceae
' Lophosoria

* Loxsomataceae
| Loxsoma
' Loxsomopsis

* Marattiaceae
| Angiopteris
| Archangiopteris
| Christensenia
| Danaea
' Marattia

* Dryopteridaceae	(definitely not monophyletic)
|* Athyrioideae
|| Adenoderris
|| Athyriopsis
|| Athyrium
|| Cornopteris
|| Deparia
|| Diplazium
|| Dryoathyrium
|| Hemidictyum
|| Lunathyrium
|' Rhachidosorus
|* Cystopteridoideae
|| Acystopteris
|| Cystopteris
|' Gymnocarpium
|* Dryopteridoideae
|| Acrophorus
|| Arachniodes
|| Coveniella
|| Cyclodium
|| Cyrtomium
|| Didymochlaena
|| Dryopolystichum
|| Dryopteris
|| Hypoderris
|| Maxonia
|| Nothoperanema
|| Olfersia
|| Peranema
|| Phanerophlebia
|| Phanerophlebiopsis
|| Polybotrya
|| Polystichum
|| Revwattsia
|| Sorolepidium
|| Stenolepia
|' Stigmatopteris
|* Hypodematioideae
|' Hypodematium
|* Lomariopsidoideae
|| Bolbitis
|| Elaphoglossum
|| Lomagramma
|| Lomariopsis
|| Peltapteris
|| Rumohra
|| Teratophyllum
|' Thysanosoria
|* Onocleoideae
|| Matteuccia
|| Onoclea
|' Onocleopsis
|* Woodsioideae
'' Woodsia

* Marsileaceae
| Marsilea
| Pilularia
' Regnellidium

* Matoniaceae
| Matonia
' Phanerosorus

* Metaxyaceae
' Metaxya

* Nephrolepidaceae
' Nephrolepis

* Ophioglossaceae
| Botrychium
| Cheiroglossum
| Helminthostachys
' Ophioglossum

* Osmundaceae
| Leptopteris
| Osmunda
' Todea

* Plagiogyriaceae
' Plagiogyria

* Polypodiaceae	(definitely not monophyletic)
|* Drynarioideae
|| Aglaomorpha
|| Arthromeris
|| Belvisia
|| Christiopteris
|| Colysis
|| Dictymia
|| Drymotaenium
|| Drynaria
|| Lecanopteris
|| Lemmaphyllum
|| Lepisorus
|| Leptochilus
|| Microsorum
|| Paraselliguea
|| Podosorus
|' Selliguea
|* Loxogrammoideae
|| Anarthropteris
|' Loxogramme
|* Oleandroideae
|| Arthropteris
|| Oleandra
|' Psammiosorus
|* Pleurosoriopsidoideae
|' Pleurosoriopsis
|* Polypodioideae
|| Campyloneurum
|| Dicranoglossum
|| Goniophlebium
|| Marginariopsis
|| Microgramma
|| Microphlebodium
|| Neurodium
|| Niphidium
|| Pecluma
|| Phlebodium
|| Platycerium
|| Pleopeltis
|| Polypodiodes
|| Polypodiopteris
|| Polypodium
|| Pseudocolysis
|| Pyrrosia
|| Solanopteris
|' Thylacopteris
|* Tectarioideae
|| Aenigmopteris
|| Atalopteris
|| Chlamydogramme
|| Ctenitis
|| Ctenitopsis
|| Cyclopeltis
|| Dictyoxiphium
|| Dryopsis
|| Heterogonium
|| Lastreopsis
|| Megalastrum
|| Pleocnemia
|| Psomiocarpa
|| Pteridrys
|| Tectaria
|| Tectaridium
'' Triplophyllum

* Psilotaceae
| Psilotum
' Tmesipteris

* Pteridaceae	(definitely not monophyletic)
|* Acrostichoideae
|| Acrostichum
|' Ceratopteris
|* Adiantoideae
|' Adiantum
|* Bommerioideae
|' Bommeria
|* Cheilanthoideae
|| Aspidotis
|| Astrolepis
|| Cheilanthes
|| Cheiloplecton
|| Negripteris
|' Pellaea
|* Coniogrammoideae
|' Coniogramme
|* Llaveoideae
|' Llavea
|* Notholaenoideae
|| Adiantopsis
|| Argyrochosma
|| Chrysochosma
|| Cryptogramma
|| Doryopteris
|| Hemionitis
|| Notholaena
|| Parahemionitis
|' Trachypteris
|* Pteridoideae
|| Actiniopteris
|| Afropteris
|| Anogramma
|| Anopteris
|| Austrogramme
|| Cerosora
|| Eriosorus
|| Jamesonia
|| Nephopteris
|| Neurocallis
|| Ochropteris
|| Onychium
|| Pentagramma
|| Pityrogramma
|| Platyzoma
|| Pteris
|| Pterozonium
|| Rheopteris
|' Syngramma
|* Taenitidoideae
'' Taenitis

* Salviniaceae
' Salvinia

* Schizaeaceae
| Actinostachys
| Anemia
| Lygodium
| Mohria
' Schizaea

* Thelypteridaceae
| Amauropelta
| Ampelopteris
| Amphineuron
| Chingia
| Christella
| Coryphopteris
| Craspedosorus
| Cyclogramma
| Cyclosorus
| Glaphyropteridopsis
| Goniopteris
| Macrothelypteris
| Meniscium
| Menisorus
| Mesophlebion
| Metathelypteris
| Nannothelypteris
| Oreopteris
| Parathelypteris
| Phegopteris
| Plesioneuron
| Pneumatopteris
| Pronephrium
| Pseudocyclosorus
| Pseudophegopteris
| Sphaerostephanos
| Stegnogramma
| Steiropteris
| Thelypteris
' Trigonospora

* Vittariaceae
| Ananthacorus
| Anetium
| Antrophyum
| Haplopteris
| Hecistopteris
| Monogramma
| Polytaenium
| Radiovittaria
| Scoliosorus
' Vittaria